Alan Townsend
Professor

Ph.D., Stanford University, 1994

INSTAAR RL1-116 (lab)
303-492-6865
alan.townsend@colorado.edu

Lab website

Alan  Townsend
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Research Interests Research Interests: Terrestrial ecosystem ecology and biogeochemistry. Research interests include the effects of human activity on regional to global scale carbon and nitrogen cycling; feedbacks between tropical ecosystems and several components of global environmental change, including climate, land-use change, and increasing nitrogen deposition; controls over nutrient limitation in moist tropical ecosystems.
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Selected Publications

Townsend, A.R., C.C. Cleveland, B.Z. Houlton, C.B. Alden and J.W.C. White. 2011. Multi-element regulation of the tropical forest carbon cycle. Frontiers in Ecology and the Environment, 1(9):9-17

Taylor, P. T. and A.R. Townsend. 2010. Stoichiometric controls over carbon-nitrate relationships from soils to the sea. Nature, 464:1178-1181.

Townsend, A.R. and R.W. Howarth. 2010. Fixing the Nitrogen Problem. Scientific American, Feb 2010, pp 64-71.

Cleveland, C.C., W.Wieder, S.C. Reed and A. R. Townsend. 2010. Experimental dry-down in a wet tropical forest increases CO2 emissions to the atmosphere. Ecology, 91: 2313-2323.

Vitousek, P.M., R. Naylor, T. Crews, M. David, L. Drinkwater, E. Holland, J. Katzenberger, L. Martinelli, P. Matson, G. Nziguheba, D. Ojima, C. Palm, G.P. Robertson, P. Sanchez, A.R. Townsend, F.S. Zhang. Nutrient imbalances along trajectories of agricultural development. 2009. Science, 324:1519-1520.

Townsend, A.R., G.P. Asner and C.C. Cleveland. 2008. The biogeochemical heterogeneity of tropical forests. Trends in Ecology and Evolution, 23(8):424-431.

Galloway, JN; Townsend, AR; Erisman, JW; Bekunda, M; Cai, ZC; Freney, JR; Martinelli, LA; Seitzinger, SP; Sutton, MA. 2008. Transformation of the nitrogen cycle: Recent trends, questions, and potential solutions. Science 320 (5878): 889-892.

Reed, S.C., C.C. Cleveland and A.R. Townsend. 2008. Tree species control rates of free-living nitrogen fixation in a lowland tropical forest. Ecology, 89(10): 2924-2934

McKenzie, V. and A.R. Townsend. Parasitic and Infectious Disease Responses to a Changing Global Nitrogen Cycle. 2007. EcoHealth, 4(4):384-396.

Townsend, A.R., C.C. Cleveland, G.P.Asner and M.M.C. Bustamante. 2007. Controls over foliar N:P ratios in tropical forests. Ecology, 88(1): 107-118.

Cleveland, C. C. and A. R. Townsend. 2006. Nitrogen and phosphorus additions cause substantial losses of soil carbon from a lowland tropical rain forest. Proceedings of the National Academy of Sciences, 103(27):10,316-10,321.

Palmer, M.A., E. Bernhardt, E. Chornesky, S.L. Collins, A. Dobson, C. Duke, B. Gold, R. Jacobson, S. Kingsland, R. Kranz, M. Mappin, M. L. Martinez, F. Micheli, J.L. Morse, M. Pace, M. Pascual, S. Palumbi, J, Reichman, W.H. Schlesinger, A. Townsend, and M. G. Turner. (alphabetical after first). 2004. Ecology for a Crowded Planet. Science 304:1251-1253.